Keep up to date here with recently published scientific research on ferns and lycophytes.
This page is a new feature on our website. Each entry has a link to the abstract and in some cases the complete article, and the list will be updated regularly to include papers as they are published.
Older articles will be put into the archive


Zhong Y, Liu Y, Wu W, Chen J, Sun C, Liu H, Shu J, Ebihara A, Yan Y, Zhou R, Schneider H.
Genomic insights into genetic diploidization in the homosporous fern Adiantum nelumboides.
Genome Biol Evol. 2022 Aug 10;. [Epub ahead of print] PMID: 35946426

Fujiwara T, Egashira T, Gutiérrez-Ortega JS, Hori K, Ebihara A, Watano Y.
Establishment of an allotetraploid fern species, Lepisorus yamaokae Seriz., between two highly niche-differentiated parental species.
Am J Bot. 2022 Aug 8;. [Epub ahead of print] PMID: 35938973

Ni R, Niu M, Fu J, Tan H, Zhu TT, Zhang J, Lou HX, Zhang P, Li JX, Cheng AX.
Molecular and structural characterization of a promiscuous chalcone synthase from the fern species Stenoloma chusanum.
J Integr Plant Biol. 2022 Aug 3;. [Epub ahead of print] PMID: 35920566

Ke BF, Wang GJ, Labiak PH, Rouhan G, Chen CW, Shepherd LD, Ohlsen DJ, Renner MAM, Karol KG, Li FW, Kuo LY.
Systematics and Plastome Evolution in Schizaeaceae.
Front Plant Sci. 2022;13:885501.
PMID: 35909781

Pelosi JA, Kim EH, Barbazuk WB, Sessa EB.
Phylotranscriptomics Illuminates the Placement of Whole Genome Duplications and Gene Retention in Ferns.
Front Plant Sci. 2022;13:882441.
PMID: 35909764

Gómez-Noguez F, Domínguez-Ugalde C, Flores-Galván C, León-Rossano LM, García BP, Mendoza-Ruiz A, Rosas-Pérez I, Mehltreter K.
Terminal velocity of fern and lycopod spores is affected more by mass and ornamentation than by size.
Am J Bot. 2022 Jul 28;. [Epub ahead of print] PMID: 35903036

Putzier CC, Polich SB, Verhoeven AS.
Sustained zeaxanthin-dependent thermal dissipation is induced by desiccation and low temperatures in the fern Polypodium virginianum.
Physiol Plant. 2022 Jun 30;:e13743. [Epub ahead of print] PMID: 35773786

Fouracre JP, Harrison CJ.
How was apical growth regulated in the ancestral land plant? Insights from the development of non-seed plants.
Plant Physiol. 2022 Jun 30;. [Epub ahead of print] PMID: 35771646

Xia ZQ, Wei ZY, Shen H, Shu JP, Wang T, Gu YF, Jaisi A, Yan YH.
Lycophyte transcriptomes reveal two whole-genome duplications in Lycopodiaceae: Insights into the polyploidization of Phlegmariurus.
Plant Divers. 2022 May;44(3):262-270.
PMID: 3576959

Yu JH, Zhang R, Liu QL, Wang FG, Yu XL, Dai XL, Liu YB, Yan YH.
Ceratopteris chunii and Ceratopteris chingii (Pteridaceae), two new diploid species from China, based on morphological, cytological, and molecular data.
Plant Divers. 2022 May;44(3):300-307.
PMID: 35769586

Putzier CC, Polich SB, Verhoeven AS.
Sustained zeaxanthin-dependent thermal dissipation is induced by desiccation and low temperatures in the fern Polypodium virginianum.
Physiol Plant. 2022 Jun 30;:e13743. [Epub ahead of print] PMID: 35773786

Fouracre JP, Harrison CJ.
How was apical growth regulated in the ancestral land plant? Insights from the development of non-seed plants.
Plant Physiol. 2022 Jun 30;. [Epub ahead of print] PMID: 35771646

Xia ZQ, Wei ZY, Shen H, Shu JP, Wang T, Gu YF, Jaisi A, Yan YH.
Lycophyte transcriptomes reveal two whole-genome duplications in Lycopodiaceae: Insights into the polyploidization of Phlegmariurus.
Plant Divers. 2022 May;44(3):262-270.
PMID: 3576959

Yu JH, Zhang R, Liu QL, Wang FG, Yu XL, Dai XL, Liu YB, Yan YH.
Ceratopteris chunii and Ceratopteris chingii (Pteridaceae), two new diploid species from China, based on morphological, cytological, and molecular data.
Plant Divers. 2022 May;44(3):300-307.
PMID: 35769586

Zheng B, Xing K, Zhang J, Liu H, Ali K, Li W, Bai Q, Ren H.
Evolutionary Analysis and Functional Identification of Ancient Brassinosteroid Receptors in Ceratopteris richardii.
Int J Mol Sci. 2022 Jun 18;23(12).
PMID: 35743240

Cai Z, Xie Z, Wang X, Zhang S, Wu Q, Yu X, Guo Y, Gao S, Zhang Y, Xu S, Wang H, Luo J.
Excavation of Genes Responsive to Brassinosteroids by Transcriptome Sequencing in Adiantum flabellulatum Gametophytes.
Genes (Basel). 2022 Jun 14;13(6).
PMID: 35741824

Matzen SL, Olson AL, Pallud CE.
Soil texture and climate limit cultivation of the arsenic hyperaccumulator Pteris vittata for phytoextraction in a long-term field study.
J Hazard Mater. 2022 Aug 15;436:129151.
PMID: 35739697

Shu JP, Wang H, Shen H, Wang RJ, Fu Q, Wang YD, Jiao YN, Yan YH.
Phylogenomic Analysis Reconstructed the Order Matoniales from Paleopolyploidy Veil.
Plants (Basel). 2022 Jun 7;11(12).
PMID: 35736680

Wan X, Zeng W, Zhang D, Wang L, Lei M, Chen T.
Changes in the concentration, distribution, and speciation of arsenic in the hyperaccumulator Pteris vittata at different growth stages.
Sci Total Environ. 2022 Jun 16;:156708. [Epub ahead of print] PMID: 35718183

Kitaoku Y, Taira T, Numata T, Ohnuma T, Fukamizo T.
Structure, mechanism, and phylogeny of LysM-chitinase conjugates specifically found in fern plants.
Plant Sci. 2022 May 6;321:111310. [Epub ahead of print] PMID: 3569691

Mehltreter K, Wachter H, Trabi C, Testo W, Sundue M, Jansen S.
Hydathodes in ferns: their phylogenetic distribution, structure, and function.
Ann Bot. 2022 Jun 13;. [Epub ahead of print] PMID: 35696156

Sun J, Li GS.
Identification of genes differentially expressed between prostrate shoots and erect shoots in the lycophyte Selaginella nipponica using an RNA-seq approach.
AoB Plants. 2022 Jun;14(3):plac018.
PMID: 35694642

Elasbali AM, Al-Soud WA, Al-Oanzi ZH, Qanash H, Alharbi B, Binsaleh NK, Alreshidi M, Patel M, Adnan M.
Cytotoxic Activity, Cell Cycle Inhibition, and Apoptosis-Inducing Potential of Athyrium hohenackerianum (Lady Fern) with Its Phytochemical Profiling.
Evid Based Complement Alternat Med. 2022;2022:2055773.
PMID: 35692581

Dai Z.
Study on the Protective Effect and Mechanism of the Rhizoma Drynariae-Epimedium Formula on Osteoarthritis in Rats.
Contrast Media Mol Imaging. 2022;2022:2869707.
PMID: 35685668

Elasbali AM, Al-Soud WA, Al-Oanzi ZH, Qanash H, Alharbi B, Binsaleh NK, Alreshidi M, Patel M, Adnan M.
Cytotoxic Activity, Cell Cycle Inhibition, and Apoptosis-Inducing Potential of Athyrium hohenackerianum (Lady Fern) with Its Phytochemical Profiling.
Evid Based Complement Alternat Med. 2022;2022:2055773.
PMID: 35692581

Maldonado I, Moreno EG, Vilca FZ.
Application of duckweed (Lemna sp.) and water fern (Azolla sp.) in the removal of pharmaceutical residues in water: State of art focus on antibiotics.
Sci Total Environ. 2022 Jun 8;:156565. [Epub ahead of print] PMID: 35690203

Carlson AB, Mathesius CA, Ballou S, Fallers MN, Gunderson TA, Hession A, Mirsky H, Stolte B, Zhang J, Woods RM, Herman RA, Roper JM.
Safety assessment of the insecticidal protein IPD079Ea from the fern, Ophioglossum pendulum.
Food Chem Toxicol. 2022 Jun 7;:113187. [Epub ahead of print] PMID: 35688270

Qoms MS, Arulrajah B, Shamsudin R, Wan Ibadullah WZ, Saari N.
Valorisation of green biomass Azolla pinnata fern: multi-parameter evaluation of processing conditions on protein extractability and their influence on the physicochemical, structural, techno-functional properties and protein quality.
J Sci Food Agric. 2022 Jun 10;. [Epub ahead of print] PMID: 35686494

Mondal S, Moktan S.
Study on the morpho-anatomy of Lepisorus species through light microscopy and scanning electron microscopy and its systematic implications.
Microsc Res Tech. 2022 Jun 9;. [Epub ahead of print] PMID: 35678524

Chao YS, Ebihara A, Chiou WL, Tsai JM, Huang YW, Ranker TA.
Reticulate evolution in the Pteris fauriei group (Pteridaceae).
Sci Rep. 2022 Jun 1;12(1):9145.
PMID: 35650209

Zheng X, Lin Z, Lu J, Ye R, Qu M, Wang J, Xu G, Ying Z, Chen S.
De novo transcriptome analysis reveals the molecular regulatory mechanism underlying the response to excess nitrogen in Azolla spp.
Aquat Toxicol. 2022 May 19;248:106202. [Epub ahead of print] PMID: 35623198

Xiang QP, Tang JY, Yu JG, Smith DR, Zhu YM, Wang YR, Kang JS, Yang J, Zhang XC.
Evolution of extremely diverged plastomes in Selaginellaceae (Lycophyte) is driven by repeat patterns and the underlying DNA maintenance machinery.
Plant J. 2022 Jun 1;. [Epub ahead of print] PMID: 35648423

Qian H, Kessler M, Vetaas OR.
Pteridophyte species richness in the central Himalaya is limited by cold climate extremes at high elevations and rainfall seasonality at low elevations.
Ecol Evol. 2022 May;12(5):e8958.
PMID: 35646313

Du XY, Kuo LY, Zuo ZY, Li DZ, Lu JM.
Structural Variation of Plastomes Provides Key Insight Into the Deep Phylogeny of Ferns.
Front Plant Sci. 2022;13:862772.
PMID: 35645990

Boeckman CJ, Ballou S, Gunderson T, Huang E, Linderblood C, Olson T, Stolte B, LeRoy K, Walker C, Wang Y, Woods R, Zhang J.
Characterization of the Spectrum of Activity of IPD079Ea: A Protein Derived From Ophioglossum pendulum (Ophioglossales: Ophioglossaceae) With Activity Against Western Corn Rootworm [Diabrotica virgifera virgifera (Coleoptera: Chrysomelidae)].
J Econ Entomol. 2022 May 29;. [Epub ahead of print] PMID: 35640234

Lu Z, Yin L, Li W, Jiang HS.
Low Concentrations of Silver Nanoparticles Inhibit Spore Germination and Disturb Gender Differentiation of Ceratopteris thalictroides (L.) Brongn.
Nanomaterials (Basel). 2022 May 18;12(10).
PMID: 35630950

Chumsuwan N, Khongkow P, Kaewsuwan S, Kanokwiroon K.
Interruptin C, a Radioprotective Agent, Derived from Cyclosorus terminans Protect Normal Breast MCF-10A and Human Keratinocyte HaCaT Cells against Radiation-Induced Damage.
Molecules. 2022 May 20;27(10).
PMID: 35630775

Tao J, Huang Z, Wang Y, Liu Y, Zhao T, Wang Y, Tian L, Cheng G.
Ethanolic Extract from Pteris wallichiana Alleviates DSS-Induced Intestinal Inflammation and Intestinal Barrier Dysfunction by Inhibiting the TLR4/NF-κB Pathway and Regulating Tight Junction Proteins.
Molecules. 2022 May 11;27(10).
PMID: 35630568

Parrish AN, Lange I, Šamec D, Lange BM.
Differential Accumulation of Metabolites and Transcripts Related to Flavonoid, Styrylpyrone, and Galactolipid Biosynthesis in Equisetum Species and Tissue Types.
Metabolites. 2022 Apr 29;12(5).
PMID: 35629907

Song C, Guan Y, Zhang D, Tang X, Chang Y.
Integrated mRNA and miRNA Transcriptome Analysis Suggests a Regulatory Network for UV-B-Controlled Terpenoid Synthesis in Fragrant Woodfern (Dryopteris fragrans).
Int J Mol Sci. 2022 May 20;23(10).
PMID: 35628519

Zhang L, Zhang LB.
Phylogeny, character evolution, and systematics of the fern family Ophioglossaceae based on Sanger sequence data, plastomes, and morphology.
Mol Phylogenet Evol. 2022 May 17;:107512. [Epub ahead of print] PMID: 35595007

Pham VT, Li SH, Dong SY.
Tectariadanangensis (Tectariaceae), a new fern species from Vietnam.
PhytoKeys. 2022;194:1-13.
PMID: 35586319

Bloesch Z, Nauheimer L, Elias Almeida T, Crayn D, Raymond Field A.
HybPhaser identifies hybrid evolution in Australian Thelypteridaceae.
Mol Phylogenet Evol. 2022 May 13;:107526. [Epub ahead of print] PMID: 35577295

Yang NY, Jia XL, Sui CX, Shen SY, Dai XL, Xue JS, Yang ZN.
Documenting the Sporangium Development of the Polypodiales Fern Pteris multifida.
Front Plant Sci. 2022;13:878693.
PMID: 35574127

Windham MD, Huiet L, Metzgar JS, Ranker TA, Yatskievych G, Haufler CH, Pryer KM.
Once more unto the breach, dear friends: Resolving the origins and relationships of the Pellaea wrightiana hybrid complex.
Am J Bot. 2022 May 14;. [Epub ahead of print] PMID: 35568966

Alejo-Jacuinde G, Herrera-Estrella L.
Exploring the High Variability of Vegetative Desiccation Tolerance in Pteridophytes.
Plants (Basel). 2022 Apr 30;11(9).
PMID: 35567223

Xia Z, Liu L, Wei Z, Wang F, Shen H, Yan Y.
Analysis of Comparative Transcriptome and Positively Selected Genes Reveal Adaptive Evolution in Leaf-Less and Root-Less Whisk Ferns.
Plants (Basel). 2022 Apr 28;11(9).
PMID: 3556719

Sugawara K, Ichio K, Ichikawa Y, Ogawa H, Suzuki S.
Effects of Pyrolysis Temperature and Chemical Modification on the Adsorption of Cd and As(V) by Biochar Derived from Pteris vittata.
Int J Environ Res Public Health. 2022 Apr 25;19(9).
PMID: 35564620

Rybczyński JJ, Kaźmierczak A, Dos Santos Szewczyk K, Tomaszewicz W, Miazga-Karska M, Mikuła A.
Biotechnology of the Tree Fern Cyathea smithii (J.D. Hooker; Soft Tree Fern, Katote) II Cell Suspension Culture: Focusing on Structure and Physiology in the Presence of 2,4-D and BAP.
Cells. 2022 Apr 20;11(9).
PMID: 35563701

Yang C, Han N, Inoue C, Yang YL, Nojiri H, Ho YN, Chien MF.
Rhizospheric plant-microbe synergistic interactions achieve efficient arsenic phytoextraction by Pteris vittata.
J Hazard Mater. 2022 Apr 6;434:128870. [Epub ahead of print] PMID: 35452977

Zuo ZY, Zhao T, Du XY, Xiong Y, Lu JM, Li DZ.
A revision of Dryopteris sect. Diclisodon (Dryopteridaceae) based on morphological and molecular evidence with description of a new species.
Plant Divers. 2022 Mar;44(2):181-190.
PMID: 35505984

Kösesakal T, Seyhan M.
Phenanthrene stress response and phytoremediation potential of free-floating fern Azolla filiculoides Lam.
Int J Phytoremediation. 2022 May 2;:1-14. [Epub ahead of print] PMID: 35501688

Wei Z, Xia Z, Shu J, Shang H, Maxwell SJ, Chen L, Zhou X, Xi W, Adjie B, Yuan Q, Cao J, Yan Y.
Corrigendum: Phylogeny and Taxonomy on Cryptic Species of Forked Ferns of Asia.
Front Plant Sci. 2022;13:888725.
PMID: 35498656

Krieg CP, Chambers SM.
The ecology and physiology of fern gametophytes: A methodological synthesis.
Appl Plant Sci. 2022 Mar-Apr;10(2):e11464.
PMID: 35495196

Nitta JH, Chambers SM.
Identifying cryptic fern gametophytes using DNA barcoding: A review.
Appl Plant Sci. 2022 Mar-Apr;10(2):e11465.
PMID: 35495195

Wu YH, Ke YT, Chan YY, Wang GJ, Kuo LY.
Integrating tissue-direct PCR into genetic identification: An upgraded molecular ecology approach to survey fern gametophytes in the field.
Appl Plant Sci. 2022 Mar-Apr;10(2):e11462.
PMID: 35495191

Hornych O, Černochová L, Lisner A, Ekrt L.
An experimental assessment of competitive interactions between sexual and apomictic fern gametophytes using Easy Leaf Area.
Appl Plant Sci. 2022 Mar-Apr;10(2):e11466.
PMID: 35495190

Schneller JJ, Farrar DR.
Photographic analysis of field-monitored fern gametophyte development and response to environmental stress.
Appl Plant Sci. 2022 Mar-Apr;10(2):e11470.
PMID: 35495189

Huang X, Wang W, Gong T, Wickell D, Kuo LY, Zhang X, Wen J, Kim H, Lu F, Zhao H, Chen S, Li H, Wu W, Yu C, Chen S, Fan W, Chen S, Bao X, Li L, Zhang D, Jiang L, Yan X, Liao Z, Zhou G, Guo Y, Ralph J, Sederoff RR, Wei H, Zhu P, Li FW, Ming R, Li Q.
The flying spider-monkey tree fern genome provides insights into fern evolution and arborescence.
Nat Plants. 2022 May 9;. [Epub ahead of print] PMID: 35534720

Zheng SQ, Song GQ, Yin CP, Chen YF, Wang SS, Shen ZB.
[A new phloroglucinol from Dryopteris fragrans and its antibacterial activity in vitro].
Zhongguo Zhong Yao Za Zhi. 2022 May;47(9):2474-2479. Chinese.
PMID: 35531695

Wang FG, Wang AH, Bai CK, Jin DM, Nie LY, Harris AJ, Che L, Wang JJ, Li SY, Xu L, Shen H, Gu YF, Shang H, Duan L, Zhang XC, Chen HF, Yan YH.
Genome size evolution of the extant lycophytes and ferns.
Plant Divers. 2022 Mar;44(2):141-152.
PMID: 35505989

Qian H, Zhang J, Jiang MC.
Global patterns of fern species diversity: An evaluation of fern data in GBIF.
Plant Divers. 2022 Mar;44(2):135-140.
PMID: 35505988

Quinlan A, Lee PH, Tang TY, Huang YM, Chiou WL, Kuo LY.
Providing the missing links in fern life history: Insights from a phenological survey of the gametophyte stage.
Appl Plant Sci. 2022 Mar-Apr;10(2):e11473.
PMID: 35495188

Farrar DR, Johnson CL.
Methodologies for soil extraction and conservation analysis of ferns and lycophytes with belowground gametophytes.
Appl Plant Sci. 2022 Mar-Apr;10(2):e11469.
PMID: 35495186

Chen KH, Xie QY, Chang CC, Kuo LY.
Mycobiome detection from a single subterranean gametophyte using metabarcoding techniques.
Appl Plant Sci. 2022 Mar-Apr;10(2):e11461.
PMID: 35495185

Suissa JS, Friedman WE.
Rapid diversification of vascular architecture underlies the Carboniferous fern radiation.
Proc Biol Sci. 2022 Apr 27;289(1973):20212209.
PMID: 35473384

Wu X, Yan A, Yang X, Banks JA, Zhang S, Zhou Y.
Cell growth dynamics in two types of apical meristems in fern gametophytes.
Plant J. 2022 Apr 21;. [Epub ahead of print] PMID: 35451138

Satjarak A, Graham LE, Trest MT, Arancibia-Avila P.
Plastid Genome of Equisetum xylochaetum from the Atacama Desert, Chile and the Relationships of Equisetum Based on Frequently Used Plastid Genes and Network Analysis.
Plants (Basel). 2022 Apr 6;11(7).
PMID: 35406981

Ben-Menni Schuler S, Hamza H, Blanca G, Romero-García AT, Suárez-Santiago VN.
Phylogeographical Analyses of a Relict Fern of Palaeotropical Flora (Vandenboschia speciosa): Distribution and Diversity Model in Relation to the Geological and Climate Events of the Late Miocene and Early Pliocene.
Plants (Basel). 2022 Mar 22;11(7).
PMID: 35406819

Kohda YH, Endo G, Kitajima N, Sugawara K, Chien MF, Inoue C, Miyauchi K.
Arsenic uptake by Pteris vittata in a subarctic arsenic-contaminated agricultural field in Japan: An 8-year study.
Sci Total Environ. 2022 Mar 25;:154830. [Epub ahead of print] PMID: 35346712

Luthardt L, Merbitz M, Fridland E, Rößler R.
Upside-down in volcanic ash: crown reconstruction of the early Permian seed fern Medullosa stellata with attached foliated fronds.
PeerJ. 2022;10:e13051.
PMID: 35341054

Saleem A, Mubeen A, Akhtar MF, Zeb A.
Polystichum braunii ameliorates airway inflammation by attenuation of inflammatory and oxidative stress biomarkers, and pulmonary edema by elevation of aquaporins in ovalbumin-induced allergic asthmatic mice.
Inflammopharmacology. 2022 Mar 7;. [Epub ahead of print] PMID: 35257281

Alejo-Jacuinde G, Kean-Galeno T, Martínez-Gallardo N, Tejero-Díez JD, Mehltreter K, Délano-Frier JP, Oliver MJ, Simpson J, Herrera-Estrella L.
Viability markers for determination of desiccation tolerance and critical stages during dehydration in Selaginella species.
J Exp Bot. 2022 Mar 21;. [Epub ahead of print] PMID: 35312760

Kinosian SP, Wolf PG.
The Natural History of Model Organisms: The biology of C. richardii as a tool to understand plant evolution.
Elife. 2022 Mar 21;11. [Epub ahead of print] PMID: 35311640

Irfan M, Jan G, Murad W, Jan FG.
Taxonomic importance of spore morphology of selected taxa of Asplenium (Aspleniaceae) from Pakistan.
Microsc Res Tech. 2022 Mar 18;. [Epub ahead of print] PMID: 35302690

Cai Z, Xie Z, Huang L, Wang Z, Pan M, Yu X, Xu S, Luo J.
Full-length transcriptome analysis of Adiantum flabellulatum gametophyte.
PeerJ. 2022;10:e13079.
PMID: 35287343

Elsebaie EM, Asker GA, Mousa MM, Kassem MM, Essa RY.
Technological and Sensory Aspects of Macaroni with Free or Encapsulated Azolla Fern Powder.
Foods. 2022 Feb 27;11(5).
PMID: 35267340

Liang SQ, Zhang XC, Wei R.
Ecological adaptation shaped the genetic structure of homoploid ferns against strong dispersal capacity.
Mol Ecol. 2022 Mar 7;. [Epub ahead of print] PMID: 35253951

da Silva GC, de Oliveira AM, Costa WK, da Silva Filho AF, Pitta MGDR, Rêgo MJBM, Antônia de Souza I, Paiva PMG, Napoleão TH.
Antibacterial and antitumor activities of a lectin-rich preparation from Microgramma vacciniifolia rhizome.
Curr Res Pharmacol Drug Discov. 2022;3:100093.
PMID: 3524333

Li Y, Wang Z, You ZH, Li LP, Hu X.
Predicting Protein-Protein Interactions via Random Ferns with Evolutionary Matrix Representation.
Comput Math Methods Med. 2022;2022:7191684.
PMID: 35242211

Han N, Yang C, Shimomura S, Inoue C, Chien MF.
Empirical Evidence of Arsenite Oxidase Gene as an Indicator Accounting for Arsenic Phytoextraction by Pteris vittata.
Int J Environ Res Public Health. 2022 Feb 4;19(3).
PMID: 35162818

Watkins JE, Testo WL.
Close observation of a common fern challenges long-held notions of how plants move. A commentary on ‘Fern fronds that move like pine cones: humidity-driven motion of fertile leaflets governs the timing of spore dispersal in a widespread fern species’.
Ann Bot. 2022 Feb 25;. [Epub ahead of print] PMID: 35211726

Hong Y, Wang Z, Li M, Su Y, Wang T.
First Multi-Organ Full-Length Transcriptome of Tree Fern Alsophila spinulosa Highlights the Stress-Resistant and Light-Adapted Genes.
Front Genet. 2021;12:784546.
PMID: 35186007

Viana JL, Dalling JW.
Soil fertility and water availability effects on trait dispersion and phylogenetic relatedness of tropical terrestrial ferns.
Oecologia. 2022 Feb 18;. [Epub ahead of print] PMID: 35179630

Andreola A, Rosini DN, Campos ML, Biasi JP, Citadini-Zanette V, da Costa Bortoluzzi RL, Miquelutti DJ, Nicoleite ER.
Ferns and lycophytes in coal mining waste and tailing landfills.
Environ Sci Pollut Res Int. 2022 Feb 15;. [Epub ahead of print] PMID: 35167024

Berim A, Gang DR.
Accumulation of Salicylic Acid and Related Metabolites in Selaginella moellendorffii.
Plants (Basel). 2022 Feb 8;11(3).
PMID: 35161442

Tomaszewicz W, Cioć M, Dos Santos Szewczyk K, Grzyb M, Pietrzak W, Pawłowska B, Mikuła A.
Enhancing In Vitro Production of the Tree Fern Cyathea delgadii and Modifying Secondary Metabolite Profiles by LED Lighting.
Cells. 2022 Jan 30;11(3).
PMID: 35159295

Elias RB, Pereira F.
New records of the critically endangered fern Grammitisazorica (Polypodiaceae).
Biodivers Data J. 2022;10:e77948.
PMID: 35136362

Hornych O, Férová A, Hori K, Košnar J, Ekrt L.
Apomictic fern fathers: An experimental approach to the reproductive characteristics of sexual, apomict and hybrid fern gametophytes.
Am J Bot. 2022 Jan 24;. [Epub ahead of print] PMID: 35072270

Wu YH, Sun CY, Ebihara A, Lu NT, Rouhan G, Kuo LY.
Two new species in the fern genus Lomariopsis (Lomariopsidaceae) from East Asia.
PhytoKeys. 2021;187:161-176.
PMID: 35068973

Chatterjee S, Bhattacharya S, Choudhury PR, Rahaman A, Sarkar A, Talukdar AD, Mandal DP, Bhattacharjee S.
Drynaria quercifolia suppresses paracetamol‑induced hepatotoxicity in mice by inducing Nrf-2.
Bratisl Lek Listy. 2022;123(2):110-119.
PMID: 3506558

Gafur NA, Sakakibara M, Komatsu S, Sano S, Sera K.
Environmental Survey of the Distribution and Metal Contents of Pteris vittata in Arsenic-Lead-Mercury-Contaminated Gold Mining Areas along the Bone River in Gorontalo Province, Indonesia.
Int J Environ Res Public Health. 2022 Jan 4;19(1).
PMID: 35010788

Zhou XM, Zhao J, Yang JJ, Le Péchon T, Zhang L, He ZR, Zhang LB.
Plastome structure, evolution, and phylogeny of Selaginella.
Mol Phylogenet Evol. 2022 Jan 11;:107410. [Epub ahead of print] PMID: 35031459

Yan H, Xu W, Zhang T, Feng L, Liu R, Wang L, Wu L, Zhang H, Zhang X, Li T, Peng Z, Jin C, Yu Y, Ping J, Ma M, He Z.
Characterization of a novel arsenite long-distance transporter from arsenic hyperaccumulator fern Pteris vittata.
New Phytol. 2022 Jan 11;. [Epub ahead of print] PMID: 35015902

Wei ZY, Gu YF, Xia ZQ, Chen LJ, Wang T, Zhang SZ, Zhao GH, Chen JB, Cao JG, Yan YH.
Dipterisshenzhenensis, a new endangered species of Dipteridaceae from Shenzhen, southern China.
PhytoKeys. 2021;186:111-120.
PMID: 35002359

Rodríguez-Pelayo C, Ambrose BA, Vasco A, Alzate JF, Pabón-Mora N.
Evolution and expression of LEAFY genes in ferns and lycophytes.
Evodevo. 2022 Jan 8;13(1):2.
PMID: 34998428

Wang R, Zhao G, Wang Z, Lin Z, Jiang X, Lin S, Chan H, Hong Y, Zhang Y, Su Y, Wang T.
The complete chloroplast genome of Vandenboschia striata, a common and widespread filmy fern (Hymenophyllaceae).
Mitochondrial DNA B Resour. 2022;7(1):128-129.
PMID: 34993335

Brilli F, Dani KGS, Pasqualini S, Costarelli A, Cannavò S, Paolocci F, Chini Zittelli G, Mugnai G, Baraldi R, Loreto F.
Exposure to different light intensities affects emission of volatiles and accumulations of both pigments and phenolics in Azolla filiculoides.
Physiol Plant. 2022 Jan 6;:e13619. [Epub ahead of print] PMID: 34988977


Wei Z, Xia Z, Shu J, Shang H, Maxwell SJ, Chen L, Zhou X, Xi W, Adjie B, Yuan Q, Cao J, Yan Y.
Phylogeny and Taxonomy on Cryptic Species of Forked Ferns of Asia.
Front Plant Sci. 2021;12:748562.
PMID: 34975938

Zhao DP, Li DL, Zhang YH, Cui Y, Xu HD, Yang B, Lei X, Zhang N.
[Total flavonoids of Drynariae Rhizoma regulates ER-p38 MAPK signaling pathway to improve scopolamine-induced learning and memory impairments in model mice].
Zhongguo Zhong Yao Za Zhi. 2021 Nov;46(22):5922-5929. Chinese.
PMID: 34951183

Kuo LY, Tang SK, Kao TT, Ebihara A, Fawcett S, Hsiao MC, Shinohara W, Dauphin B.
A dormant resource for genome size estimation in ferns: C-value inference of the Ophioglossaceae using herbarium specimen spores.
Appl Plant Sci. 2021 Nov-Dec;9(11-12):e11452.
PMID: 34938613

Yadav BL, Meghvansi MK, Meena K, Gena CB.
Discovery of a new species of Adder’s tongue fern from India with comparative analysis of morphological and molecular attributes.
Sci Rep. 2021 Dec 22;11(1):24396.
PMID: 3493786

Irfan M, Jan G, Murad W, Jan FG, Rauf A, Alsayari A, Almarhoon ZM, Mabkhot YN.
Ethnomedicinal and traditional uses of the Ferns of Khyber Pakhtunkhwa, Pakistan.
Braz J Biol. 2021;84:e250256.
PMID: 34932624

Yang FC, Sarathchandra C, Liu JX, Huang HP, Gou JY, Li Y, Mao XY, Wen HT, Zhao J, Yang MF, Homya S, Prueksakorn K.
How fern and fern allies respond to heterogeneous habitat – a case in Yuanjiang dry-hot valley.
Commun Integr Biol. 2021;14(1):248-260.
PMID: 34925688

Suissa JS.
Fern fronds that move like pine cones: humidity-driven motion of fertile leaflets governs the timing of spore dispersal in a widespread fern species.
Ann Bot. 2021 Nov 19;. [Epub ahead of print] PMID: 34878516

Fernández H, Grossmann J, Gagliardini V, Feito I, Rivera A, Rodríguez L, Quintanilla LG, Quesada V, Cañal MJ, Grossniklaus U.
Sexual and Apogamous Species of Woodferns Show Different Protein and Phytohormone Profiles.
Front Plant Sci. 2021;12:718932.
PMID: 34868105

Melnikov D, Tretyakova A, Grudanov N, Baranova O, Senator S, Muldashev A, Podgaevskaya E, Zolotareva N, Erokhin N, Vorobiev A, Knyazev MS, Glazunov V, Kapitonova O, Allayarova V, Naumenko N, Efimik E, Malykh S, Merker V, Morozyuk Y, Burundukova D, Shubin D, Shilov D.
Pteridophyte distribution of the Urals and adjacent areas: a dataset.
Biodivers Data J. 2021;9:e76680.
PMID: 34866963

Wei R, Yang J, He LJ, Liu HM, Hu JY, Liang SQ, Wei XP, Zhao CF, Zhang XC.
Plastid phylogenomics provides novel insights into the infrafamilial relationship of Polypodiaceae.
Cladistics. 2021 Dec;37(6):717-727.
PMID: 34841589

Sciandrello S, Cambria S, Del Galdo GG, Tavilla G, Minissale P.
Unexpected Discovery of Thelypteris palustris (Thelypteridaceae) in Sicily (Italy): Morphological, Ecological Analysis and Habitat Characterization.
Plants (Basel). 2021 Nov 12;10(11).
PMID: 34834810

Ben-Menni Schuler S, Picazo-Aragonés J, Rumsey FJ, Romero-García AT, Suárez-Santiago VN.
Macaronesia Acts as a Museum of Genetic Diversity of Relict Ferns: The Case of Diplazium caudatum (Athyriaceae).
Plants (Basel). 2021 Nov 10;10(11).
PMID: 34834788

Huang K, Shang H, Zhou Q, Wang Y, Shen H, Yan Y.
Volatiles Induced from Hypolepis punctata (Dennstaedtiaceae) by Herbivores Attract Sclomina erinacea (Hemiptera: Reduviidae): Clear Evidence of Indirect Defense in Fern.
Insects. 2021 Oct 28;12(11).
PMID: 3482177

Shu JP, Liu ZY, Gu ZR, Chen LJ, Wei HJ, Zhou XL, Yan YH, Wang RJ.
Dryopteriswulingshanensis (Dryopteridaceae), a new species from Hunan, China.
PhytoKeys. 2021;185:17-26.
PMID: 34819778

Trujillo-Argueta S, Del Castillo RF, Tejero-Diez D, Matias-Cervantes CA, Velasco-Murguía A.
DNA barcoding ferns in an unexplored tropical montane cloud forest area of southeast Oaxaca, Mexico.
Sci Rep. 2021 Nov 24;11(1):22837.
PMID: 34819549

Colpo A, Baldisserotto C, Pancaldi S, Sabia A, Ferroni L.
Photosystem II photoinhibition and photoprotection in a lycophyte, Selaginella martensii.
Physiol Plant. 2021 Nov 22;:e13604. [Epub ahead of print] PMID: 34811759

Matzen SL, Lobo GP, Fakra SC, Kakouridis A, Nico PS, Pallud CE.
Arsenic hyperaccumulator Pteris vittata shows reduced biomass in soils with high arsenic and low nutrient availability, leading to increased arsenic leaching from soil.
Sci Total Environ. 2021 Nov 19;:151803. [Epub ahead of print] PMID: 34808151

Medeiros-Fonseca B, Abreu-Silva AL, Medeiros R, Oliveira PA, Gil da Costa RM.
Pteridium spp. and Bovine Papillomavirus: Partners in Cancer.
Front Vet Sci. 2021;8:758720.
PMID: 34796228

Tanaka M, Koeduka T, Matsui K.
Green Leaf Volatile-Burst in Selaginella moellendorffii.
Front Plant Sci. 2021;12:731694.
PMID: 34777416

Zhang L, Rothfels CJ, Ebihara A, Schuettpelz E, Le Péchon T, Kamau P, He H, Zhou XM, Prado J, Field A, Yatskievych G, Gao XF, Zhang LB.
A global plastid phylogeny of the brake fern genus Pteris (Pteridaceae) and related genera in the Pteridoideae.
Cladistics. 2015 Aug;31(4):406-423.
PMID: 34772268

López CL, Mayta C, Naoki K, Quezada JAN, Hensen I, Gallegos SC.
Bracken fern does not diminish arbuscular mycorrhizal fungus inoculum potential in tropical deforested areas.
Mycorrhiza. 2021 Nov 8;. [Epub ahead of print] PMID: 34750710

Zhang D, Yang W, Liu H, Cui Y, Wang L, Liu X, Zhang G.
The complete chloroplast genome of a distinctive fern, Coniogramme intermedia Hieron. (Pteridaceae).
Mitochondrial DNA B Resour. 2021;6(12):3318-3319.
PMID: 34746400

Costarelli A, Cannavò S, Cerri M, Pellegrino RM, Reale L, Paolocci F, Pasqualini S.
Light and Temperature Shape the Phenylpropanoid Profile of Azolla filiculoides Fronds.
Front Plant Sci. 2021;12:727667.
PMID: 34745161

Fan XP, Thi Lu N, Li CX, Knapp R, He H, Zhou XM, Wan X, Zhang L, Gao XF, Zhang LB.
Phylogeny, biogeography, and character evolution in the fern family Hypodematiaceae.
Mol Phylogenet Evol. 2021 Nov 1;:107340. [Epub ahead of print] PMID: 34737000

Wickell D, Kuo LY, Yang HP, Dhabalia Ashok A, Irisarri I, Dadras A, de Vries S, de Vries J, Huang YM, Li Z, Barker MS, Hartwick NT, Michael TP, Li FW.
Underwater CAM photosynthesis elucidated by Isoetes genome.
Nat Commun. 2021 Nov 3;12(1):6348.
PMID: 34732722

Klaus KV, Schulz C, Bauer DS, Stützel T.
Historical biogeography of the ancient lycophyte genus Selaginella: early adaptation to xeric habitats on Pangea.
Cladistics. 2017 Oct;33(5):469-480.
PMID: 34724754

Matongera TN, Mutanga O, Sibanda M.
Characterizing bracken fern phenological cycle using time series data derived from Sentinel-2 satellite sensor.
PLoS One. 2021;16(10):e0257196.
PMID: 34710104

Rencoret J, Gutiérrez A, Marques G, Del Río JC, Tobimatsu Y, Lam PY, Pérez-Boada M, Ruiz-Dueñas FJ, Barrasa JM, Martínez AT.
New Insights on Structures Forming the Lignin-Like Fractions of Ancestral Plants.
Front Plant Sci. 2021;12:740923.
PMID: 34691117

Suissa JS, Kinosian SP, Schafran PW, Bolin JF, Carl Taylor W, Zimmer EA.
Homoploid hybrids, allopolyploids, and high ploidy levels characterize the evolutionary history of a western North American quillwort complex (Isoëtes).
Mol Phylogenet Evol. 2021 Oct 20;:107332. [Epub ahead of print] PMID: 34687842

Zemanová V, Pavlíková D, Hnilička F, Pavlík M.
Arsenic Toxicity-Induced Physiological and Metabolic Changes in the Shoots of Pteris cretica and Spinacia oleracea.
Plants (Basel). 2021 Sep 25;10(10).
PMID: 34685818

Yang L, Huang Y, Lima LV, Sun Z, Liu M, Wang J, Liu N, Ren H.
Corrigendum: Rethinking the Ecosystem Functions of Dicranopteris, a Widespread Genus of Ferns.
Front Plant Sci. 2021;12:770556.
PMID: 34675959

Fang T, Motte H, Parizot B, Beeckman T.
Early “Rootprints” of Plant Terrestrialization: Selaginella Root Development Sheds Light on Root Evolution in Vascular Plants.
Front Plant Sci. 2021;12:735514.
PMID: 34671375

Durieux T, Lopez MA, Bronson AW, Tomescu AMF.
A new phylogeny of the cladoxylopsid plexus: contribution of an early cladoxylopsid from the Lower Devonian (Emsian) of Quebec.
Am J Bot. 2021 Oct 19;. [Epub ahead of print] PMID: 34664712

Matia-Merino L, Goh KKT, Singh H.
A natural shear-thickening water-soluble polymer from the fronds of the black tree fern, Cyathea medullaris: Influence of salt, pH and temperature.
Carbohydr Polym. 2012 Jan 4;87(1):131-138.
PMID: 34662941

Iqbal MS, Ahmad KS, Ali MA, Akbar M, Mehmood A, Nawaz F, Hussain SA, Arshad N, Munir S, Arshad H, Shahbaz K, Bussmann RW.
An ethnobotanical study of wetland flora of Head Maralla Punjab Pakistan.
PLoS One. 2021;16(10):e0258167.
PMID: 34648500

Zhang Y, Li L, Friml J.
Evaluation of Gravitropism in Non-seed Plants.
Methods Mol Biol. 2022;2368:43-51.
PMID: 34647246

Mehltreter K, Tenhaken R, Jansen S.
Nectaries in ferns: their taxonomic distribution, structure, function, and sugar composition.
Am J Bot. 2021 Oct 13;. [Epub ahead of print] PMID: 34643269

Al-Musawi TJ, Mengelizadeh N, Taghavi M, Mohebi S, Balarak D.
Activated carbon derived from Azolla filiculoides fern: a high-adsorption-capacity adsorbent for residual ampicillin in pharmaceutical wastewater.
Biomass Convers Biorefin. 2021 Oct 3;:1-13. [Epub ahead of print] PMID: 34631357

Lehtonen S, Poczai P, Sablok G, Hyvönen J, Karger DN, Flores J.
Exploring the phylogeny of the marattialean ferns.
Cladistics. 2020 Dec;36(6):569-593.
PMID: 34618987

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